I need help configuring the A* Pathfinding Project to suit my level. Is it even possible?

Try using the AILerp Component instead of your AIPathAlignedToSurface Component and see if that works. It should follow the path exacly.

Idk what’s going on when gravity is disabled. Maybe the “Continue to Exact Destination” somehow triggers right at the beginning? Or the “Closest On Node Surface” from the Seeker aauses the path to have issues on recalculation?

You can also try to disable recalculation for these tests. Right now it’s important to get something working and then you can try to change things up but you can also always go back to the one version that’s working.